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{{Taxobox
| color = pink
| name = Chimpanzees<ref name=MSW3>{{MSW3 Groves|pages=182-183}}</ref>
| image = South Djoum Chimp.jpg
| image_width = 250px
| image_caption = [[Common Chimpanzee]]<br>in [[Cameroon]]'s [[South Province, Cameroon|South Province]]
| regnum = [[Animal]]ia
| phylum = [[Chordata]]
| classis = [[Mammal]]ia
| ordo = [[Primate]]s
| familia = [[Hominidae]]
| subfamilia = [[Homininae]]
| tribus = [[Hominini]]
| subtribus = '''Panina'''
| genus = '''''Pan'''''
| genus_authority = [[Lorenz Oken|Oken]], 1816
| type_species = ''[[Common Chimpanzee|Simia troglodytes]]''
| type_species_authority = [[Johann Friedrich Blumenbach|Blumenbach]], 1775
| range_map = Pan.svg
| range_map_width = 250px
| range_map_caption = distribution of ''Pan spp.''
| subdivision_ranks = [[Species]]
| subdivision =
''[[Common Chimpanzee|Pan troglodytes]]''<br/>
''[[Bonobo|Pan paniscus]]''
}}
Chimpanzee, often shortened to chimp, is the common name for the two extant species in the genus Pan. The better known chimpanzee is Pan troglodytes, the Common Chimpanzee, living primarily in West, and Central Africa. Its cousin, the Bonobo or "Pygmy Chimpanzee" as it is known archaically, Pan paniscus, is found in the forests of the Democratic Republic of the Congo. The Congo River forms the boundary between the two species.
Most adherents to the theory of evolution hold the view that chimpanzees and humans shared a close common ancestor.
==Measurements==
A full grown adult male chimpanzee can weigh from 35-70 kilograms (75-155 pounds) and stand 0.9-1.2 meters (3-4 feet) tall, while females usually weigh 26-50 kg (57-110 pounds) and stand 0.66-1 meters (2.0-3.5 feet) tall.
==Lifespan==
Chimpanzees rarely live past the age of 40 in the wild, but have been known to reach the age of 60 in captivity, and ''[[Tarzan]]'' star [[Cheeta]] is still alive [[as of 2006]] at the age of 74.
==Chimpanzee differences==
[[Image:Bonobo_009.jpg|thumb|Bonobo]]
Anatomical differences between Common and Pygmy Chimpanzees are slight, but in [[sex|sexual]] and social behaviour there are marked differences. Common Chimpanzees have an [[omnivore|omnivorous]] [[diet (nutrition)|diet]], a troop [[predation|hunting]] culture based on beta males led by an [[alpha male]], and highly complex social relationships; [[bonobo|Bonobos]], on the other hand, have a mostly [[herbivorous]] diet and an [[egalitarian]], [[matriarchal]], sexually receptive behavior. The exposed skin of the face, hands and feet varies from pink to very dark in both species, but is generally lighter in younger individuals, darkening as maturity is reached. Bonobos have longer arms and tend to walk upright more often than the Common Chimpanzee.
==Taxonomic relationships==
[[Image:Hominoid taxonomy 7.png|thumb|350px|The taxonomic relationships of Hominoidea]]
The genus ''Pan'' is now considered to be part of the subfamily [[Homininae]] to which [[human]]s also belong. [[Biology|Biologists]] believe that the two species of chimpanzees are the closest living [[evolution|evolutionary]] relatives to [[human]]s. It is thought that humans shared a common ancestor with chimpanzees and gorillas as recently as [[Pliocene|four]] to [[Miocene|seven million years ago]]. Groundbreaking research by [[Mary-Claire King]] in 1973 found 99% identical [[DNA]] between human beings and chimpanzees,<ref>Mary-Claire King, ''Protein polymorphisms in chimpanzee and human evolution'', Doctoral dissertation, University of California, Berkeley (1973).</ref> although research since has modified that finding to about 94%<ref name=ns>{{cite web | url = http://www.sciam.com/article.cfm?chanID=sa003&articleID=9D0DAC2B-E7F2-99DF-3AA795436FEF8039 |date = 2006-12-19 | title = Humans and Chimps: Close But Not That Close | publisher = Scientific American | accessdate = 2006-12-20}}</ref> commonality, with at least some of the difference occurring in 'junk' DNA. It has even been proposed that ''troglodytes'' and ''paniscus'' belong with ''sapiens'' in the genus ''[[Homo (genus)|Homo]]'', rather than in ''Pan''. One argument for this is that other species have been reclassified to belong to the same genus on the basis of less genetic similarity than that between humans and chimpanzees. Indeed cladistic taxonomy, based on both genetic difference and date of likely divergence, is very clear in placing both extant species of ''Pan'' in the genus ''Homo'', mainly because the genus ''Homo'' takes precedence on account of being coined first. It is very important, however, to consider where the differences in the genome appear.
A study published by Clark and Nielsen of [[Cornell University]] in the December 2003 issue of the journal ''[[Science (journal)|Science]]'' highlights differences related to one of humankind's defining qualities — the ability to understand [[language]] and to communicate through speech. These macro-phenotypic differences, however, may owe less to physiology than might be assumed given that ''Homo sapiens'' developed modern cultural features long after the modern physiological features were in place and indeed competed averagely against other species of ''Homo'' with regard to tools, etc for many millennia. Differences also exist in the genes for smell, in genes that regulate the metabolism of amino acids and in genes that may affect the ability to digest various proteins. See the [[ape#History of hominoid taxonomy|history of hominoid taxonomy]] for more about the history of the classification of chimpanzees. See [[Human evolutionary genetics]] for more information on the speciation of humans and great apes.
===Fossils===
Many [[Human evolution|human fossils]] have been found, but chimpanzee fossils were not described until 2005. Existing chimpanzee populations in West and Central Africa do not overlap with the major human fossil sites in East Africa. However, chimpanzee fossils have now been reported from [[Kenya]]. This would indicate that both humans and members of the ''Pan'' [[clade]] were present in the East African [[Rift Valley]] during the Middle [[Pleistocene]].<ref name=firstfossil>{{cite journal | title = First fossil chimpanzee | last = McBrearty | first = S. | coauthors = N. G. Jablonski | journal = [[Nature (journal)|Nature]] | date = 2005-09-01 | volume = 437 | pages = 105-108 | id = {{Entrez Pubmed|16136135}}}}</ref>
| color = pink
| name = Chimpanzees<ref name=MSW3>{{MSW3 Groves|pages=182-183}}</ref>
| image = South Djoum Chimp.jpg
| image_width = 250px
| image_caption = [[Common Chimpanzee]]<br>in [[Cameroon]]'s [[South Province, Cameroon|South Province]]
| regnum = [[Animal]]ia
| phylum = [[Chordata]]
| classis = [[Mammal]]ia
| ordo = [[Primate]]s
| familia = [[Hominidae]]
| subfamilia = [[Homininae]]
| tribus = [[Hominini]]
| subtribus = '''Panina'''
| genus = '''''Pan'''''
| genus_authority = [[Lorenz Oken|Oken]], 1816
| type_species = ''[[Common Chimpanzee|Simia troglodytes]]''
| type_species_authority = [[Johann Friedrich Blumenbach|Blumenbach]], 1775
| range_map = Pan.svg
| range_map_width = 250px
| range_map_caption = distribution of ''Pan spp.''
| subdivision_ranks = [[Species]]
| subdivision =
''[[Common Chimpanzee|Pan troglodytes]]''<br/>
''[[Bonobo|Pan paniscus]]''
}}
Chimpanzee, often shortened to chimp, is the common name for the two extant species in the genus Pan. The better known chimpanzee is Pan troglodytes, the Common Chimpanzee, living primarily in West, and Central Africa. Its cousin, the Bonobo or "Pygmy Chimpanzee" as it is known archaically, Pan paniscus, is found in the forests of the Democratic Republic of the Congo. The Congo River forms the boundary between the two species.
Most adherents to the theory of evolution hold the view that chimpanzees and humans shared a close common ancestor.
==Measurements==
A full grown adult male chimpanzee can weigh from 35-70 kilograms (75-155 pounds) and stand 0.9-1.2 meters (3-4 feet) tall, while females usually weigh 26-50 kg (57-110 pounds) and stand 0.66-1 meters (2.0-3.5 feet) tall.
==Lifespan==
Chimpanzees rarely live past the age of 40 in the wild, but have been known to reach the age of 60 in captivity, and ''[[Tarzan]]'' star [[Cheeta]] is still alive [[as of 2006]] at the age of 74.
==Chimpanzee differences==
[[Image:Bonobo_009.jpg|thumb|Bonobo]]
Anatomical differences between Common and Pygmy Chimpanzees are slight, but in [[sex|sexual]] and social behaviour there are marked differences. Common Chimpanzees have an [[omnivore|omnivorous]] [[diet (nutrition)|diet]], a troop [[predation|hunting]] culture based on beta males led by an [[alpha male]], and highly complex social relationships; [[bonobo|Bonobos]], on the other hand, have a mostly [[herbivorous]] diet and an [[egalitarian]], [[matriarchal]], sexually receptive behavior. The exposed skin of the face, hands and feet varies from pink to very dark in both species, but is generally lighter in younger individuals, darkening as maturity is reached. Bonobos have longer arms and tend to walk upright more often than the Common Chimpanzee.
==Taxonomic relationships==
[[Image:Hominoid taxonomy 7.png|thumb|350px|The taxonomic relationships of Hominoidea]]
The genus ''Pan'' is now considered to be part of the subfamily [[Homininae]] to which [[human]]s also belong. [[Biology|Biologists]] believe that the two species of chimpanzees are the closest living [[evolution|evolutionary]] relatives to [[human]]s. It is thought that humans shared a common ancestor with chimpanzees and gorillas as recently as [[Pliocene|four]] to [[Miocene|seven million years ago]]. Groundbreaking research by [[Mary-Claire King]] in 1973 found 99% identical [[DNA]] between human beings and chimpanzees,<ref>Mary-Claire King, ''Protein polymorphisms in chimpanzee and human evolution'', Doctoral dissertation, University of California, Berkeley (1973).</ref> although research since has modified that finding to about 94%<ref name=ns>{{cite web | url = http://www.sciam.com/article.cfm?chanID=sa003&articleID=9D0DAC2B-E7F2-99DF-3AA795436FEF8039 |date = 2006-12-19 | title = Humans and Chimps: Close But Not That Close | publisher = Scientific American | accessdate = 2006-12-20}}</ref> commonality, with at least some of the difference occurring in 'junk' DNA. It has even been proposed that ''troglodytes'' and ''paniscus'' belong with ''sapiens'' in the genus ''[[Homo (genus)|Homo]]'', rather than in ''Pan''. One argument for this is that other species have been reclassified to belong to the same genus on the basis of less genetic similarity than that between humans and chimpanzees. Indeed cladistic taxonomy, based on both genetic difference and date of likely divergence, is very clear in placing both extant species of ''Pan'' in the genus ''Homo'', mainly because the genus ''Homo'' takes precedence on account of being coined first. It is very important, however, to consider where the differences in the genome appear.
A study published by Clark and Nielsen of [[Cornell University]] in the December 2003 issue of the journal ''[[Science (journal)|Science]]'' highlights differences related to one of humankind's defining qualities — the ability to understand [[language]] and to communicate through speech. These macro-phenotypic differences, however, may owe less to physiology than might be assumed given that ''Homo sapiens'' developed modern cultural features long after the modern physiological features were in place and indeed competed averagely against other species of ''Homo'' with regard to tools, etc for many millennia. Differences also exist in the genes for smell, in genes that regulate the metabolism of amino acids and in genes that may affect the ability to digest various proteins. See the [[ape#History of hominoid taxonomy|history of hominoid taxonomy]] for more about the history of the classification of chimpanzees. See [[Human evolutionary genetics]] for more information on the speciation of humans and great apes.
===Fossils===
Many [[Human evolution|human fossils]] have been found, but chimpanzee fossils were not described until 2005. Existing chimpanzee populations in West and Central Africa do not overlap with the major human fossil sites in East Africa. However, chimpanzee fossils have now been reported from [[Kenya]]. This would indicate that both humans and members of the ''Pan'' [[clade]] were present in the East African [[Rift Valley]] during the Middle [[Pleistocene]].<ref name=firstfossil>{{cite journal | title = First fossil chimpanzee | last = McBrearty | first = S. | coauthors = N. G. Jablonski | journal = [[Nature (journal)|Nature]] | date = 2005-09-01 | volume = 437 | pages = 105-108 | id = {{Entrez Pubmed|16136135}}}}</ref>