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J.B.S. Haldane (1892-1964) was a famous evolutionist and geneticist.  He was one of the three founders of “population genetics” as a field of study.  In 1957 Haldane published a paper outlining a serious problem for the theory of evolution.  He called this problem the “cost of substitution” but is has become known as “Haldane’s Dilemna”.

'''J.B.S. Haldane''' (1892-1964) was a famous evolutionist] and geneticist.  He was one of the three founders of “population genetics” as a field of study.  In 1957 Haldane published a paper outlining a serious problem for the [[theory of evolution]].  He called this problem the “cost of substitution” but is has become known as “Haldane’s Dilemna”.

For evolution to occur, a beneficial mutation must not only occur, but it must be copied to the point where the mutated genome substitutes for the un-mutated genome in the population.  This substitution is limited by the reproduction rate of the species in question.

For evolution to occur, a beneficial mutation must not only occur, but it must be copied to the point where the mutated genome substitutes for the un-mutated genome in the population.  This substitution is limited by the reproduction rate of the species in question.